Silver Perch
Silver perch Bidyanus bidyanus, is a freshwater
finfish species exotic to Western Australia, and has been imported
into this State since 1950 for the purpose of stocking inland farm
dams. Imports have only been allowed under license, subject to disease-free
certification and on the basis that the fish were being placed into
impounded or confined waters. Lawrence (1995) discussed the prospects
for aquaculture of silver perch in Western Australia and since then
the industry has only expanded slowly. A similar pattern existed
in the early days of the silver perch industry
in NSW, where rapid progress is now being made, and this species
should still be regarded as having significant potential in WA.
However, its limited salinity tolerance may restrict its usage for
highly saline inland waters. This publication updates the information
provided by Lawrence (1995).
Silver Perch research
Research into the hatchery production of silver perch commenced
in New South Wales more than 20 years ago at the Narrandera Fisheries
Centre. Since 1990, silver perch has received increasing attention
as a candidate finfish species for aquaculture in NSW, Queensland
and WA. In 1994, a silver perch hatchery began operation
in Parkerville, WA. Silver perch fingerlings are now produced at several
farms and on-grown commercially at a number of locations including
Northampton, Jurien, Pemberton, Esperance, Dandaragan, Dardanup and
the Perth metropolitan area.
Silver Perch distribution
Silver perch is native to the extensive Murray-Darling
river system of south-eastern Australia. Significant populations,
with patchy distributions, of silver perch are found in NSW and southern
Queensland. Silver perch also occurs in small numbers in the colder
regions of Victoria. Over the past 50 years, the distribution and
abundance of silver perch has decreased (dramatically in the last
ten years) with silver perch given the conservation status of 'potentially
threatened' (Jackson, 1994) and it is possible that the species will
become 'endangered' in the future (Rowland & Bryant, 1995).
Silver Perch distribution within Western Australia
Although silver perch is not native to WA, it has been brought here
for both recreational and aquaculture ventures. Further distribution
of this species within WA however, raises a number of important issues
relating to the transfer of fish to waters outside their natural or
previous distribution (called 'translocation'). These
include the potential of the species to establish feral populations
in the wild and to introduce disease. As a result the Department of
Fisheries has developed a policy to enable the continued development
of a silver perch industry in WA in an environmentally acceptable
manner. The
Department of Fisheries Management Paper No. 145 outlines areas
within the State where silver perch may or may not be farmed and
how silver perch can be sold or imported into WA.
The areas in which silver perch are permitted are determined by
the conservation value of drainage basins:
Category 1: Drainage basins, or areas within drainage
basins in which silver perch farms and stocking are not permitted;
Category 2: Drainage basins in which silver perch
farms and stocking may be permitted, subject to conditions; and
Category 3: Drainage basins in which silver perch
farms and stocking will be permitted, unless otherwise indicated
by the Executive Director of the
Department of Fisheries.
The appropriate temperature for growing this species in aquaculture
is 22oC to 28oC (Rowland, 1995c), indicating most areas below the
Tropic of Capricorn would be suited to silver perch farming. This
coincides with the current distribution of yabbies in Western Australia.
Silver Perch Habitat
Silver perch in their native east coast habitat live in rivers,
lakes and reservoirs, although they are found in waters ranging from
static to fast flowing.
Silver Perch General Biology
Silver perch is a potamodromous species, meaning
it migrates wholly within fresh water lakes and streams. In the wild,
adults undertake extensive upstream migrations and require an increase
in water level to induce spawning (Cadwallader, 1986). Spawning usually
occurs during the summer floods, when water temperatures are above
20oC and inundated flood plains provide juvenile nursery areas (Allen,
1989).
The female Silver Perch
Female silver perch reach sexual maturity at three
or four years of age and approximately 340 mm in length, while males
mature at two or three years of age and approximately 233 mm in length.
There is no sexual dimorphism within the species except for a slight
difference in body contour in near-ripe females (Lake, 1968).
Spawning behaviour is very aggressive and injuries such as scale
loss and frayed fins are sustained by both males and females. Female
silver perch usually release most of their eggs in a single spawning.
Water-hardened silver perch eggs are between 2.5 to 3.0 mm in diameter,
are non-adhesive and slightly negatively buoyant (Thurstan &
Rowland, 1995).
Silver perch can tolerate relatively poor water quality, such as
dissolved oxygen down to 2.2 mg/L; pH up to 10.2; and un-ionised
ammonia up to 0.7 mg/L, although long-term exposure to un-ionised
ammonia concentrations above 0.1 mg/L may adversely affect growth
(Rowland, 1995c). Silver perch tolerate long-term exposure to salinities
up to 5 parts per thousand (ppt) (Rowland & Bryant, 1995). However,
Guo et al. (1995) showed that if acclimated correctly, silver perch
are able to survive in salinities up to 12 ppt but die within 18
hours in salinities of 15 ppt or higher.
Culturing Silver Perch
The growing of silver perch to market size involves three distinct
phases (Rowland, 1995b):
Hatchery Phase: Most growers will not be involved with this phase.
It lasts roughly eight weeks, involves collection and spawning of
brood fish, incubation of eggs, and rearing of larvae through to
fry (30 mm total length or 0.5 g).
Fingerling Phase: Fry are grown in farmers' ponds until they are
about 15 to 20 g (three to four months at summer temperatures),
when they should be harvested and graded. Grading is particularly
important, as research has shown that fingerling size is hugely
variable from 2 to 50 grams total length after just three to four
months.
Growout Phase: Graded fingerlings are stocked in ponds for growout
to market size (10 to 15 months).
Silver Perch Hatchery Phase
Silver perch broodstock are available from several growers in WA
and should be held in earthen ponds for at least seven months prior
to the breeding season (Rowland, 1984). A good typical pond for
holding broodstock has a surface area of 0.1ha. About two-thirds
of the pond is 1 m deep, the other third around 2.4 m deep. A concrete
sump or raceway is built into the deepest section of the pond to
facilitate the recapture of the fish when the pond is drained. Silver
perch broodstock are stocked at densities up to 200/ha. Both sexes
are placed together, usually with a few extra males to ensure fertilisation
of eggs when recaptured for breeding (Thurstan & Rowland, 1995).
Silver perch broodstock are removed from ponds and are then induced
to spawn in tanks. Broodstock will mature naturally but generally
females must be induced to spawn by injection of a hormone, human
chorionic gonadotrophin (hCG) at a dose rate of 200 International
Units per kilogram of body weight (IU/kg). After hormonal injection,
ripe fish placed in tanks spawn in around 36 to 40 hours and the
eggs are fertilised naturally (MacKinnon, 1989).
In a spawning season, a large broodstock female weighing 1.8 kg
can produce approximately 500,000 eggs. Fertilised eggs are collected
from tanks using a 2 mm mesh and placed in cone-shaped fibreglass
tanks with gentle aeration at a density of up to 500 eggs per litre.
Eggs hatch in approximately 24 to 30 hours at 24oC. Larval development
takes 18 days. WA producers have had to adapt eastern states culture
techniques to WA conditions and this has resulted in slight changes
to methodology (G. Whisson, pers. comm.).
Although both silver perch eggs and juveniles are susceptible to
saline conditions, at low salinities (<5 ppt) juvenile silver
perch demonstrate a better survival rate over a 20-day period than
those hatched in fresh water. The increased survival in low salinities
may be due to the salt's inhibition of microbial pathogens (Rowland,
1984; Merrick & Schmida, 1984; Allen, 1989). This provides an
opportunity for the farming of this species in farm dams in the
wheat belt area of WA which are moderately salt-affected.
Larval-rearing ponds need to be filled 10 to 14 days before stocking,
with water pre-filtered through a 0.5 mm screen to remove potential
predators such as aquatic insects and other fish species. When full,
ponds are fertilised with inorganic and organic fertilisers to produce
phytoplankton followed by zooplankton blooms. Larvae completely
resorb their yolk sac five to six days after hatching and are now
stocked directly into ponds after hatching (Thurstan & Rowland,
1995).
Fertilisation of ponds promotes populations of micro-organisms
(bacteria, protozoans) and blooms of microscopic algae, which are
then grazed on by zooplankton (copepods, rotifers, cladocerans).
Silver perch larvae feed first on freshwater rotifers (Brachionus
spp.), followed by copepods, then cladocerans as the larvae increase
in size. Rotifer numbers need to be around five per millilitre (ml)
to support a stocking ratio of 50 to 100 larvae m2.
During this period frequent monitoring of water quality (temperature,
pH, dissolved oxygen and total ammonia nitrogen), zooplankton populations
and examination of larvae for parasites are essential (Thurstan
& Rowland, 1995c; Lambert, 1998).
Metamorphosis begins with the development of fin rays after about
seven days and is completed at around day 10 to 14 or when larvae
are 10 mm total length (TL). After five to six weeks the fish should
have full scale development and be between 25 and 35 mm TL (0.5
grams). Once larvae have metamorphosed they need to be weaned onto
artificial feed - this should begin in the third week after stocking
(S. Rowland, pers. comm.). |